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Reproduced with permission from Rasmussen, L. E. L.; Wittemyer, George. Chemosignalling of musth by individual wild African elephants (Loxodonta africana): Implications for conservation and management Proceedings of the Royal Society of London - Series B: Biological Sciences. 269(1493). 22 April, 2002. 853-860. Copyright ©2002 The Royal Society. and George Wittemyer.

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